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"Walaupun mereka agak pelbagai dari segi ekologi dan akibatnya, dalam penyesuaian kepada gaya hidup mereka yang berbeza, juga dari segi morfologi dan etologi, beberapa ciri masih menyatukan unggas air dan darat. Walau bagaimanapun, kebanyakan daripada ini adalah plesiomorph untuk Neornithes secara keseluruhan, dan juga dikongsi dengan paleognathae.[perlu rujukan]
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