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In page Phenotype:

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Phenotypes develop through an interaction of genes and their immediate cellular environment, the cellular environment being under the influence of the host-organism's interaction with its environment. Thus there is a multiplicity of ways that genes and phenotypes interact. Most simply, for example, we might say an albino phenotype develops as a consequence of a mutation in the gene encoding tyrosinase which is a key enzyme in melanin formation. Even here, however, exposure to UV radiation can increase melanin production, hence the environment plays a role in this phenotype as well. For most complex phenotypes the precise genetic mechanism remains unknown.[citation needed]