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In page Portable Executable:

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In a .NET executable, the PE code section contains a stub that invokes the CLR virtual machine startup entry, _CorExeMain or _CorDllMain in mscoree.dll, much like it was in Visual Basic executables. The virtual machine then makes use of .NET metadata present, the root of which, IMAGE_COR20_HEADER (also called "CLR header") is pointed to by IMAGE_DIRECTORY_ENTRY_COMHEADER (the entry was previously used for COM+ metadata in COM+ applications, hence the name[citation needed]) entry in the PE header's data directory. IMAGE_COR20_HEADER strongly resembles PE's optional header, essentially playing its role for the CLR loader.[1]