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In page Centriole:

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LECA, the last common ancestor of all eukaryotes was a ciliated cell with centrioles.[citation needed] Some lineages of eukaryotes, such as land plants, do not have centrioles except in their motile male gametes. Centrioles are completely absent from all cells of conifers and flowering plants, which do not have ciliate or flagellate gametes.[1] It is unclear if the last common ancestor had one[2] or two cilia.[3] Important genes such as those coding for centrins, required for centriole growth, are only found in eukaryotes, and not in bacteria or archaea.[2]