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In page Kerogen:

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Another possible method of formation is that vanabin-containing organisms cleave the core out of chlorin-based compounds such as the magnesium in chlorophyll and replace it with their vanadium center in order to attach and harvest energy via light-harvesting complexes. It is theorized that the bacteria contained in worm castings, Rhodopseudomonas palustris, do this during its photoautotrophism mode of metabolism. Over time colonies of light harvesting bacteria solidify, forming kerogen[citation needed] .