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In page Reelin:

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In 1994, a new allele of reeler was obtained by means of insertional mutagenesis.[4] This provided the first molecular marker of the locus, permitting the RELN gene to be mapped to chromosome 7q22 and subsequently cloned and identified.[5] Japanese scientists at Kochi Medical School successfully raised antibodies against normal brain extracts in reeler mice, later these antibodies were found to be specific monoclonal antibodies for reelin, and were termed CR-50 (Cajal-Retzius marker 50).[6] They noted that CR-50 reacted specifically with Cajal-Retzius neurons, whose functional role was unknown until then.[citation needed]