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In page Manorialism:

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Nor were manors held necessarily by lay lords rendering military service (or again, cash in lieu) to their superior: a substantial share (estimated by value at 17% in England in 1086) belonged directly to the king, and a greater proportion (rather more than a quarter) were held by bishoprics and monasteries. Ecclesiastical manors tended to be larger, with a significantly greater villein area than neighbouring lay manors.[citation needed]