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In page Carrying capacity:

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This system can also be applied to natural areas. Grazing megaherbivores at roughly 1 GV/ha is considered sustainable in central European grasslands, although this varies widely depending on many factors. In ecology it is theoretically (i.e. cyclic succession, patch dynamics, Megaherbivorenhypothese) taken that a grazing pressure of 0.3 GV/ha by wildlife is enough to hinder afforestation in a natural area. Because different species have different ecological niches, with horses for example grazing short grass, cattle longer grass, and goats or deer preferring to browse shrubs, niche differentiation allows a terrain to have slightly higher carrying capacity for a mixed group of species, than it would if there were only one species involved.[citation needed]