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In page Neo-Latin:

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The digraphs ae and oe were typically written using the ligatures æ and œ (e.g. Cæsar, pœna) except when part of a word in all capitals, such as in titles, chapter headings, or captions. More rarely (and usually in 16th- to early 17th-century texts) the e caudata was used as a substitute for the digraphs.[citation needed]