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In page Rust (fungus):

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Puccinia graminis is a macrocyclic heteroecious fungus that causes wheat stem rust disease.[citation needed] The sexual stage in this fungus occurs on the alternate host – barberry – and not wheat. The durable spore type produced on the alternate host allows the disease to persist in wheat even in more inhospitable environments. Planting resistant crops will prevent disease, however, virulence mutations will give rise to new strains of fungi that overcome plant resistance.[citation needed] Although the disease cannot be stopped by removal of the alternate host, the life cycle is disrupted and the rate of evolution is decreased because of reduced genetic recombination. This allows resistance bred crops to remain effective for a longer period of time.[1][2]