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In page Adenosine diphosphate:
"During the initial phases of glycolysis and the TCA cycle, cofactors such as NAD+ donate and accept electrons[1] that aid in the electron transport chain's ability to produce a proton gradient across the inner mitochondrial membrane.[2] The ATP synthase complex exists within the mitochondrial membrane (FO portion) and protrudes into the matrix (F1 portion). The energy derived as a result of the chemical gradient is then used to synthesize ATP by coupling the reaction of inorganic phosphate to ADP in the active site of the ATP synthase enzyme; the equation for this can be written as ADP + Pi → ATP.[citation needed]
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