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In page Space Shuttle Columbia:

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Columbia was the only operational orbiter with black chines. These were added because at first, shuttle designers did not know how reentry heating would affect the craft's upper wing surfaces.[citation needed] The chines allowed Columbia to be easily recognized at a distance, unlike the subsequent orbiters. The black chines were a late modification and were a high temperature paint over the white FRSI. The chines were modified on Columbia shortly before rollover to the Vehicle Assembly Building in late 1980 for STS-1. The only other orbiter with black chines was Pathfinder,[1] but it was a cosmetic test article and only gained it when was refurbished.