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In page Stoma:
"There is little evidence of the evolution of stomata in the fossil record, but they had appeared in land plants by the middle of the Silurian period.[1] They may have evolved by the modification of conceptacles from plants' alga-like ancestors.[2] However, the evolution of stomata must have happened at the same time as the waxy cuticle was evolving – these two traits together constituted a major advantage for early terrestrial plants.[citation needed]
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