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In page Horizontal gene transfer:

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The archaeon Sulfolobus solfataricus, when UV irradiated, strongly induces the formation of type IV pili which then facilitates cellular aggregation.[3][4] Exposure to chemical agents that cause DNA damage also induces cellular aggregation.[3] Other physical stressors, such as temperature shift or pH, do not induce aggregation, suggesting that DNA damage is a specific inducer of cellular aggregation.[citation needed]