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In page Progesterone:
"The conversion of pregnenolone to progesterone takes place in two steps. First, the 3β-hydroxyl group is oxidized to a keto group and second, the double bond is moved to C4, from C5 through a keto/enol tautomerization reaction.[2] This reaction is catalyzed by 3β-hydroxysteroid dehydrogenase/δ5-4-isomerase.[citation needed]
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