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In page Aesthetics:

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Aesthetic judgment is closely tied to disgust.[citation needed] Responses like disgust show that sensory detection is linked in instinctual ways to facial expressions including physiological responses like the gag reflex. Disgust is triggered largely by dissonance; as Darwin pointed out, seeing a stripe of soup in a man's beard is disgusting even though neither soup nor beards are themselves disgusting. Aesthetic judgments may be linked to emotions or, like emotions, partially embodied in physical reactions. For example, the awe inspired by a sublime landscape might physically manifest with an increased heart-rate or pupil dilation.