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In page Tilde:

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Later, it was used to make abbreviations in medieval Latin documents. When an ⟨n⟩ or ⟨m⟩ followed a vowel, it was often omitted, and a tilde (physically, a small ⟨N⟩) was placed over the preceding vowel to indicate the missing letter; this is the origin of the use of tilde to indicate nasalization (compare the development of the umlaut as an abbreviation of ⟨e⟩.)[citation needed] A tilde represented an omitted ⟨a⟩ or a syllable containing it.[1] The practice of using the tilde over a vowel to indicate omission of an ⟨n⟩ or ⟨m⟩ continued in printed books in French as a means of reducing text length until the 17th century. It was also used in Portuguese and Spanish.[citation needed]