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In page Sense of balance:

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Since the cupula is part of the bony labyrinth, it rotates along with actual head movement, and by itself without the endolymph, it cannot be stimulated and therefore, could not detect movement. Endolymph follows the rotation of the canal; however, due to inertia its movement initially lags behind that of the bony labyrinth. The delayed movement of the endolymph bends and activates the cupula. When the cupula bends, the connected stereocilia bend along with it, activating chemical reactions in the hair cells surrounding crista ampullaris and eventually create action potentials carried by the vestibular nerve signaling to the body that it has moved in space.[citation needed]