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Two extinct species were named from the fossil record.[1] Pandion homalopteron described by Stuart L. Warter in 1976 was found in marine Middle Miocene deposits of the Barstovian age in the southern part of California. The second species Pandion lovensis was described by Jonathan J. Becker in 1985 and found in Florida; it dates to the Late Clarendonian and possibly represents a separate lineage from that of P. homalopteron and P. haliaetus. A number of claw fossils have been recovered from Pliocene and Pleistocene sediments in Florida and South Carolina.[citation needed]